VOLCANOES NATIONAL PARK, Rwanda -- Deep in Rwanda's steep-sloped forest, the mountain gorillas look both endearing and intimidating. A tourist might feel conflicting impulses to shy away and reach for a hug (the latter is not advised) when a gorilla brushes past on a path. The way a gorilla snoozes, scratches a leg or casts an inquiring glance -- it all seems familiar, and yet wild.
"You can't tell what they're thinking," said John Scott, a retired chemical engineer from Britain's Worcester area who trekked to the high-altitude habitat to see the creatures with close genetic links to humans.
This sense of kinship helps explain why increasing numbers of tourists are heading to Rwanda's Volcanoes National Park, fueling an industry seen as key to the welfare of the critically endangered subspecies as well as the national economy. Those visitors can also be a threat because gorillas are vulnerable to human diseases and so reduced in numbers that a veterinary team called Gorilla Doctors cares for sick and injured apes.
The mountain gorilla population dropped sharply in the last century because of poaching, illness and human encroachment, although the numbers are now rising. These days, an estimated 900 mountain gorillas live in Rwanda and neighbouring Congo and Uganda.
In Rwanda, conservation is big business. Eighty individual permits to see the gorillas for one hour are available daily for a maximum price of $750 each, and 20 per cent of permit revenue goes to schools, clinics and other local community projects, the park website says.
More than 20,000 people visited Rwanda's gorillas in 2014, nearly three times as many as in 2003, according to government figures. Many came from the United States, Britain, Australia, Germany and Canada. Last week, people in hiking gear sipped coffee and milled around at the park headquarters before breaking into groups of eight, the limit for tourist parties visiting separate gorilla families in the dense undergrowth.
"How's the pace? Are you feeling the mountains yet or not?" guide Ferdinand Ndamiyabo asked his group, including an Associated Press team, during a hike up a volcano that is home to a family of gorillas called Amahoro, which means "peace" in the Rwandan language. It took close to two hours of walking to reach the gorillas, a relatively gentle climb in mild weather through tangled vines, stinging nettles and other lush vegetation.
Ndamiyabo earlier laid out rules for the encounter: Don't point, speak softly, don't cough or sneeze in the animals' direction and stay a minimum of 23 feet (7 metres) away. If a gorilla approaches, crouch down, don't make eye contact and make a low sound similar to that of clearing the throat, which gorillas use to express friendliness.
What awaited in a clearing were drowsy gorillas, including two young ones that idly grappled and another that groomed Gahinga, an adult male silverback that dominates the Amahoro. Gahinga eventually rolled off his back and rested his great head on an arm, watching the camera-toting arrivals. He made a low sound.
"The silverback is saying, 'No problem, my friends, take as many pictures as you want,"' Ndamiyabo declared.
Dr. Jean Bosco Noheli, a Gorilla Doctors veterinarian who accompanied the tourists, noted a wrist wound on Karisimbi, a female gorilla named after the highest volcano in the border-spanning Virunga mountain range where gorillas live. He described the injury as superficial and said there was no need for doctors to intervene, a complex process that would require darting the gorilla with a tranquilizer and likely fending off other gorillas before treatment on the spot.
Another member of the group is Kajoriti, a male who lost a hand to a poacher's snare.
American researcher Dian Fossey, who won the confidence of gorillas by imitating their noises, moving on her knuckles and chewing on vegetation, brought international attention to the primates' plight. Fossey, whose book "Gorillas in the Mist" inspired a movie starring Sigourney Weaver, was murdered at her Rwandan research camp in 1985 and is buried at a mountain gravesite.
Rwanda descended into bloody chaos during its 1994 genocide, and tourism only returned to Volcanoes National Park at the end of the decade. Since then, Microsoft co-founder Bill Gates and Hollywood actors are among those who visited Rwanda's mountain gorillas, whose home is a two-hour drive from Kigali, the capital.
The group led by guide Ndamiyabo followed the gorillas after their morning nap. At times, gorillas fell in behind the startled tourists, almost bumping into them as they advanced.
Sarah Scott, a nurse and wife of tourist John Scott, said the close encounter was awe-inspiring.
The gorillas seemed so human -- whether "grooming or passing wind" -- but also huge and powerful, she said, adding: "One swipe of the hand and that's it."